(IP (NP-SBJ (NN 蝮蛇科) (NN 蛇类)) (VP (VV 具有) (NP-OBJ (CP (WHNP-1 (-NONE- *OP*)) (CP (IP (NP-SBJ (-NONE- *T*-1)) (VP (VV 分泌) (NP-OBJ (NN 毒素)))) (DEC 的))) (NP (NN 毒腺)))))
ARG0: *T*-1 ARG1: 毒素 REL: 分泌